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In C#, is there a difference between default(Nullable<long>); (or default(long?);) and default(long); ?

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Well yes. The default value of a nullable or other reference type is null while the default value for a long or other value type is 0 (and any other members set to their defaults).

In this case:

default(Nullable<long>) == null
default(long?) == null

default(long) == 0L